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Such_Saturation
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This is also relevant and assures that I'm not "making this up". Acknowledging Selection At Sub-organismal Levels Resolves Controversy On Pro-cooperation MechanismsVery interesting thread. This gem of a post is hiding just below and very applicable.
Cooperation, Not Competition, Drives Evolution
selection can operate on organismal and sub-organismal 'entities' such that partner fidelity feedback at sub-organismal level can appear as partner choice at organismal level.